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    Molecular phylogeny confirms the subspecies delineation of the Malayan Siamang (Symphalangus syndactylus continentis) and the Sumatran Siamang (Symphalangus syndactylus syndactylus) based on the hypervariable region of mitochondrial DNA
    “… subspecies were proposed as early as 1908 by Thomas namely Symphalangus s. syndactylus and Symphalangus…”
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